| Boulenophrys | |
|---|---|
 | |
| Boulenophrys baishanzuensis | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Amphibia |
| Order: | Anura |
| Family: | Megophryidae |
| Subfamily: | Megophryinae |
| Genus: | Boulenophrys Fei, Ye & Jiang, 2016 |
| Type species | |
| Boulenophrys boettgeri （Boulenger, 1899） | |
| Synonyms[1] | |
| |
Boulenophrys is a genus of frogs in the family Megophryidae. They occur in the China, Mainland Southeast Asia and Northeast India. It had been placed variously as a subgenus or synonymy of Megophrys. Dubois, Ohler and Pyron first recognized that Panophrys is preoccupied and employed Boulenophrys as the generic name rather than Tianophrys under the Principle of First Revisor.[1]
Most of the species in this genus were formerly considered species of the genus Megophrys.
Taxonomy
The following species are recognised in the genus Boulenophrys:[2]
- Boulenophrys acuta (Wang, Li & Jin, 2014)
- Boulenophrys angka (Wu, Suwannapoom, Poyarkov, Chen, Pawangkhanant, Xu, Jin, Murphy & Che, 2019)
- Boulenophrys anlongensis (Li, Lu, Liu & Wang, 2020)
- Boulenophrys baishanzuensis (Wu, Li, Liu, Wang & Wu, 2020)
- Boulenophrys baolongensis (Ye, Fei & Xie, 2007)
- Boulenophrys binchuanensis (Ye & Fei, 1995)
- Boulenophrys binlingensis (Jiang, Fei & Ye, 2009)
- Boulenophrys boettgeri (Boulenger, 1899)
- Boulenophrys brachykolos (Inger & Romer, 1961)
- Boulenophrys caobangensis (Nguyen, Pham, Nguyen, Luong & Ziegler, 2020)
- Boulenophrys caudoprocta (Shen, 1994)
- Boulenophrys cheni (Wang & Liu, 2014)
- Boulenophrys chishuiensis (Xu, Li, Liu, Wei & Wang, 2020)
- Boulenophrys congjiangensis (Luo, Wang, Wang, Lu, Wang, Deng & Zhou, 2021)
- Boulenophrys daiyunensis Lyu, Wang & Wang, 2021
- Boulenophrys dalaolingensis Wang, Chen & Liu, 2025[3]
- Boulenophrys daoji (Lyu, Zeng, Wang & Wang, 2021)
- Boulenophrys daweimontis (Rao & Yang, 1997)
- Boulenophrys daxuemontis Liu, Li, Cheng, Wei, Wang & Cheng, 2025[4]
- Boulenophrys dongguanensis (Wang & Wang, 2019)
- Boulenophrys dupanglingensis Xiao & Mo, 2025[5]
- Boulenophrys elongata Zeng, Wang, Chen, Xiao, Zhan, Li & Lin, 2024[6]
- Boulenophrys fansipanensis (Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley, 2018)
- Boulenophrys fengshunensis Wang, Zeng, Lyu, and Wang, 2022
- Boulenophrys frigida (Tapley, Cutajar, Nguyen, Portway, Mahony, Nguyen, Harding, Luong & Rowley, 2021)
- Boulenophrys gutu Kuang, Wei & Shi, 2025[7]
- Boulenophrys hengshanensis Qian, Hu, Mo, Gao, Zhang & Yang, 2023[8]
- Boulenophrys hoanglienensis (Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley, 2018)
- Boulenophrys huangniushiensis H.-T. Wang, Wu, Song, Huang, Zhuo et Y.-Y. Wang, 2025[9]
- Boulenophrys huangshanensis (Fei & Ye, 2005)
- Boulenophrys hungtai Wang, Zeng, Lyu, Xiao, and Wang, 2022
- Boulenophrys insularis (Wang, Liu, Lyu, Zeng & Wang, 2017)
- Boulenophrys jiangi (Liu, Li, Wei, Xu, Cheng, Wang & Wu, 2020)
- Boulenophrys jingdongensis (Fei & Ye, 1983)
- Boulenophrys jinggangensis (Wang, 2012)
- Boulenophrys jiulianensis (Wang, Zeng, Lyu & Wang, 2019)
- Boulenophrys kuatunensis (Pope, 1929)
- Boulenophrys leishanensis (Li, Xu, Liu, Jiang, Wei & Wang, 20192018)
- Boulenophrys liboensis (Zhang, Li, Xiao, Li, Pan, Wang, Zhang & Zhou, 2017)
- Boulenophrys lichun Lin, Chen, Li, Peng, Zeng & Wang, 2024[10]
- Boulenophrys lini (Wang & Yang, 2014)
- Boulenophrys lishuiensis (Wang, Liu & Jiang, 2017)
- Boulenophrys lushuiensis (Shi, Li, Zhu, Jiang, Jiang & Wang, 2021)
- Boulenophrys minor (Stejneger, 1926)
- Boulenophrys mirabilis (Lyu, Wang & Zhao, 2020)
- Boulenophrys mufumontana (J.Wang, Lyu & Y.Y.Wang, 2019)
- Boulenophrys nankunensis (Wang, Zeng & .Wang, 2019)
- Boulenophrys nanlingensis (Lyu, J.Wang, Liu & Y.Y.Wang, 2019)
- Boulenophrys nebulosa Shen & Li, 2025[11]
- Boulenophrys obesa (Wang, Li & Zhao, 2014)
- Boulenophrys ombrophila (Messenger & Dahn, 2019)
- Boulenophrys omeimontis (Liu, 1950)
- Boulenophrys palpebralespinosa (Bourret, 1937)
- Boulenophrys parva (Boulenger, 1893)
- Boulenophrys pepe Wang, Lin, Gan, Chen, Yu, Pan, Xiao & Zeng, 2024[12]
- Boulenophrys puningensis Wang, Zeng, Lyu, Xiao, and Wang, 2022
- Boulenophrys qianbeiensis (Su, Shi, Wu, Li, Yao, Wang & Li, 2020)
- Boulenophrys rubrimera (Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong & Rowley, 2017)
- Boulenophrys sangzhiensis (Jiang, Ye & Fei, 2008)
- Boulenophrys sanmingensis (Lyu & Wang, 2021)
- Boulenophrys shimentaina (Lyu, Liu & Wang, 2020)
- Boulenophrys shuichengensis (Tian & Sun, 1995)
- Boulenophrys shunhuangensis (Wang, Deng, Liu, Wu & Liu, 2019)
- Boulenophrys spinata (Liu & Hu, 1973)
- Boulenophrys tongboensis (Wang & Lyu, 2021)
- Boulenophrys tuberogranulata (Shen, Mo & Li, 2010)
- Boulenophrys wugongensis (J.Wang, Lyu & Y.Y.Wang, 2019)
- Boulenophrys wuliangshanensis (Ye & Fei, 1995)
- Boulenophrys wushanensis (Ye & Fei, 1995)
- Boulenophrys xiangnanensis (Lyu, Zeng & Wang, 2020)
- Boulenophrys xianjuensis (Wang, Wu, Peng, Shi, Lu & Wu, 2020)
- Boulenophrys yangchunensis Zhao, Lin, Li, Lyu, Zheng, Zeng, Borzée & Wang, 2025[13]
- Boulenophrys yangmingensis (Lyu, Zeng & Wang, 2020)
- Boulenophrys yaoshanensis Qi, Mo, Lyu, Wang & Wang, 2021
- Boulenophrys yingdeensis Qi, Lyu, Wang & Wang, 2021
- Boulenophrys yezhongensis Liu, Feng, Shen, Li, Cheng, Wei, Wang & Su, 2025[14]
- Boulenophrys youran Xiao, Shi & Mo, 2026[15]
- Boulenophrys yunkaiensis Qi, Wang, Lyu & Wang, 2021
References
- 1 2 Dubois, Alain; Ohler, Annemarie; Pyron, R. Alexander (26 February 2021). "New concepts and methods for phylogenetic taxonomy and nomenclature in zoology, exemplified by a new ranked cladonomy of recent amphibians (Lissamphibia)". Megataxa. 5 (1). doi:10.11646/megataxa.5.1.1.
- ↑ Frost, Darrel R. (2021). "Boulenophrys Fei, Ye, and Jiang, 2016, 1822". Amphibian Species of the World: An Online Reference. Version 6.1. American Museum of Natural History. Retrieved 29 July 2022.
- ↑ Ruiqi Wang, Lu Chen, Jinliang Chen, Haochen Huang, Gangzhi Peng, Honglin Peng, Xinzhang Gao, Bangqing Chen and Naiyi Liu. 2025. A new horned toad of Boulenophrys (Anura, Megophryidae) was discovered in Hubei Province, China. ZooKeys. 1253: 305-320. DOI: doi.org/10.3897/zookeys.1253.154757
- ↑ Jing Liu, Shize Li, Yanlin Cheng, Gang Wei, Bin Wang and Gang Cheng. 2025. A New Species of the Genus Boulenophrys (Anura, Megophryidae) from Southwest China. Biodiversity Data Journal. 13: e153987. DOI: doi.org/10.3897/BDJ.13.e153987
- ↑ Bei Xiao, Jiayan Xi, Shengchao Shi, Hui Li, Leqiang Zhu, Ayinuer Maimaiti, Yalan Xu, Shunhua Liao, Bin Wang and Xiaoyang Mo. 2025. A New Species of the Genus Boulenophrys (Anura, Megophryidae) from southern Hunan Province, Central China. Animals. 15(3); 440. DOI: doi.org/10.3390/ani15030440
- ↑ Zhaochi Zeng, Jian Wang, Honghui Chen, Weiwen Xiao, Binbin Zhan, Yuanhang Li and Shishi Lin. 2024. A New Species of the Genus Boulenophrys (Anura, Megophryidae) from Eastern Guangdong, China. Asian Herpetological Research. 15(1); https://www.sciengine.com/AHR/doi/10.3724/ahr.2095-0357.2023.0038
- ↑ Dai-Yong Kuang, Yi-Fu Wei, Yi-Sha Luo, Kang-Wen Pei, Ying-Yue Cao, Meng-Fei Zhang, Tai-Fu Huang, Ling Pu and Sheng-Chao Shi. 2025. A New Species of Boulenophrys (Megophyridae) from Mt. Hengshan, Hunan Province, China, with Re-Description on B. hengshanensis. Animals. 15(18), 2745. DOI: doi.org/10.3390/ani15182745 [19 September 2025]
- ↑ Boulenophrys hengshanensis Qian, Hu, Mo, Gao, Zhang & Yang, 2023
- ↑ Hao-Tian WANG, Xiao-Gang WU, Han-Ming SONG, Zhong HUANG, Xiao-Hai ZHUO and Ying-Yong WANG. 2025. A New Species of the Genus Boulenophrys (Anura: Megophryidae: Megophryinae) from Jiulianshan National Nature Reserve, southern Jiangxi, China. Zootaxa. 5691(3); 376-398. DOI: doi.org/10.11646/zootaxa.5691.3.1 [2025-09-10]
- ↑ Shi-Shi Lin, Hong-Hui Chen, Yuan-Hang Li, Zhao-Ning Peng, Zhao-Chi Zeng and Jian Wang. 2024. A New Boulenophrys Species (Anura, Megophryidae) from the coastal hills of eastern Fujian Province, China. ZooKeys. 1216: 1-15. DOI doi.org/10.3897/zookeys.1216.130017
- ↑ Deming Shen, Haiying Zhou, Chaoyang Wang, Jun Xu, Shi Xu, Yifeng Zhong, Zhenyu Wang, Hao Cai, Yahui Wu, Meicai Wei and Yankuo Li. 2025. A New Species of Boulenophrys (Anura, Megophryidae) from northern Jiangxi, China. Zoosystematics and Evolution. 101(4): 2251-2266. DOI: doi.org/10.3897/zse.101.166556
- ↑ Jian Wang, Shi-Shi Lin, Jun-Shen Gan, Hong-Hui Chen, Ling-Min Yu, Zhong Pan, Jia-Jie Xiao, Zhao-Chi Zeng. 2024. A New Species of the Genus Boulenophrys from South China (Anura, Megophryidae). Zootaxa. 5514(5); 451-468. DOI: doi.org/10.11646/zootaxa.5514.5.3
- ↑ Yi-Yang Zhao, Shi-Shi Lin, Yuan-Hang Li, Zhi-Tong Lyu, Ke Zheng, Zhao-Chi Zeng, Amaël Borzée and Jian Wang. 2025. A New Species of Chinese horned toad (Anura, Megophryidae, Boulenophrys) from western Guangdong Province, China, with recommendations for extinction risk assessments for several related congeners. Herpetozoa. 38: 345-357. DOI: doi.org/10.3897/herpetozoa.38.e171400
- ↑ Jing Liu, Chao-Bo Feng, Tuo Shen, Shi-Ze Li, Yanlin Cheng, Gang Wei, Bin Wang and Haijun Su. 2025. A New Species of the genus Boulenophrys (Anura, Megophryidae) from Guizhou, China. Herpetozoa. 38: 117-136. DOI: doi.org/10.3897/herpetozoa.38.e155430
- ↑ Bei Xiao, Leqiang Zhu, Lu Wang, Hui Li, Ayinuer Maimaiti, Shize Li, Guoai Zhang, Wuzhou Liu, Dongbo Fei, Changle Liao, Xiaoquan Tang, Shengchao Shi and Xiaoyang Mo. 2026. A New Species of the Genus Boulenophrys (Anura, Megophryidae) from Nanshan National Park, Hunan Province, China. Zoosystematics and Evolution. 102(1): 53-70. DOI: doi.org/10.3897/zse.102.175092 [05 Jan 2026]